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This article is about the organ in all vertebrate species. For other uses of the term Heart, please see Heart (disambiguation).

The heart is a muscular pump that drives the cardiovascular system. It differs in the cardiovascular systems of mammals, birds, amphibians, reptiles, and fish.


The heart is composed of cardiac muscle that differs slightly from the skeletal and smooth muscle found elsewhere in the body. This special type of muscle adjusts the rate of muscular contraction, allowing the heart to maintain a regular pumping rhythm. The main parts of the heart are the chambers, the valves, and the electrical nodes.

There are two different types of heart chambers. The first is the atrium, which receives blood returning to the heart through the veins. The right atrium pumps blood to the right ventricle, and the left atrium pumps blood into the left ventricle. This blood is then pumped from the atrium into the second chamber called the ventricle. The ventricles are much larger than the atria and their thick, muscular walls are used to forcefully pump the blood from the heart to the body and lungs (or gills).


The valves found within the heart are situated between the atria and ventricles, and also between the ventricles and major arteries. These valves are opened and closed by pressure changes within the chambers, and act as a barrier to prevent the backflow of blood. The characteristic beating heart sounds heard through a stethoscope are the result of vibrations caused by the closing of the respective valves.

There are two different electrical nodes, or groups of specialized cells, located in the cardiac tissue. The first is the sinoatrial node (SA), commonly called the pacemaker. The pacemaker is embedded in the wall of the right atrium. This small patch of tissue experiences rhythmic excitation and the impulse rapidly spreads throughout the atria, causing a muscular contraction and the pumping of blood from the atria to the ventricles.

The other node, the atrioventricular node, relays the impulse of the SA node to the ventricles. It delays the impulse to prevent the ventricles from contracting at the same time as the atria, thus giving them time to fill with blood. The cycle of contraction of the heart muscle is called a heartbeat, the rate of which varies greatly between organisms.


A blood vessel is a hollow tube for transporting blood. There are three main types of blood vessels:

These main blood vessels function to transport blood through the entire body and exchange oxygen and nutrients for carbon dioxide and wastes.

The arteries carry blood away from the heart, and are under high pressure from the pumping of the heart. To maintain their structure under this pressure, they have thick, elastic walls to allow stretch and recoil. The large pulmonary artery carries unoxygenated blood from the right ventricles to the lung, where it gives off carbon dioxide and receives oxygen. The aorta is the largest artery. It carries oxygenated blood from the left ventricle to the body. The arteries branch and eventually lead to capillary beds.

The capillaries make up a network of tiny vessels with extremely thin, highly permeable walls. They are present in all of the major tissues of the body and function in the exchange of gases, nutrients, and fluids between the blood, body tissues, and alveoli of the lungs.

Comparative anatomy

Mammals and birds

Mammalian and avian hearts have four chambers – two atria and two ventricles. This is the most efficient system, as deoxygenated and oxygenated bloods are not mixed. The right atrium receives deoxygenated blood from the body through both the inferior and superior vena cava. The blood then passes to the right ventricle to be pumped through the pulmonary arteries to the lungs, where it becomes oxygenated. It returns to the left atrium via the pulmonary veins, this oxygen-rich blood is then passed to the left ventricle and pumped through the aorta to the rest of the body. The aorta is the largest artery and has an enormous amount of stretch and elasticity to withstand the pressure created by the pumping ventricle. The four-chambered heart ensures that the tissues of the body are supplied with oxygen-saturated blood to facilitate sustained muscle movement. Also, the larger oxygen supply allows these warm-blooded organisms to achieve thermoregulation (body temperature maintenance).

Human heart

For more information, see: Human heart.

In the human body the heart is situated in the middle of the thorax and weighs about 300-350g in an adult. It consists of four chambers, the two upper atria and the two lower ventricles.

Amphibians and reptiles

Amphibians and reptiles, by contrast, have a three-chambered heart. The three-chambered heart consists of two atria and one ventricle. (The crocodile is sometimes said to have a four-chambered heart. The separation of the ventricles is not complete, however, because a hole remains in the septum (wall) that divides the two chambers.) Blood leaving the ventricle passes into one of two vessels. It either travels through the pulmonary arteries leading to the lungs or through a forked aorta leading to the rest of the body. Oxygenated blood returning to the heart from the lungs through the pulmonary vein passes into the left atrium, while deoxygenated blood returning from the body through the sinus venosus passes into the right atrium. Both atria empty into the single ventricle, mixing the oxygen-rich blood returning from the lungs with the oxygen-depleted blood from the body tissues. While this system assures that some blood always passes to the lungs and then back to the heart, the mixing of blood in the single ventricle means the organs are not getting blood saturated with oxygen. This is not as efficient as a four-chambered system, which keeps the two circuits separate, but it is sufficient for these cold-blooded organisms. The heart rate of amphibians and reptiles is very dependent upon temperature. The higher the temperature, the faster the heart beat.


Fish possess the simplest type of true heart – a two-chambered organ composed of one atrium and one ventricle. A rudimentary valve is located between the two chambers. Blood is pumped from the ventricle through the conus arteriosus to the gills. The conus arteriosus is like the aorta in other species. At the gills, the blood receives oxygen and gets rid of carbon dioxide. Blood then moves on to the organs of the body, where nutrients, gases, and wastes are exchanged. There is no division of the circulation between the gills and the body. That is, the blood travels from the heart to the gills, and then directly to the body before returning to the atrium through the sinus venosus to be circulated again. The heart rates of fish fall within the wide range of 60-240 beats per minute, depending upon species and water temperature. The fish's heart rate will be slower at lower temperatures.