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[[Image:Nwhi_-_French_Frigate_Shoals_reef_-_many_fish.jpg|thumb|300px|right|Marine life can be very abundant.]]
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{{Image|Marine-biology.gif|right|300px|Marine biologists<br><small>Photo credit: Michael Nishizaki</small>}}
'''Marine biology''' is the scientific study of the [[plant]]s, [[animal]]s and other organisms that live in the [[ocean]].  Given that in [[biology]] many [[scientific classification | phyla]], families and genera have some species that live in the [[sea]] and others that live on land, marine biology focuses on species based on their [[natural environment | environment]] rather than on [[taxonomy]].
'''Marine biology''' is the scientific study of the [[plant]]s, [[animal]]s and other organisms that live in the earth's [[ocean]], seas, and related salt-water [[Natural environment|environments]]. Marine biologists are concerned not just with individual types of animals, but also with their inter-relationships with each other and their habitat.


There are many practical reasons to study marine biology.  Marine life represents a vast resource, providing [[food]], [[medicine]], and raw materials, in addition to helping to support [[recreation]] and [[tourism]] all over the world. At a fundamental level, marine life helps determine the very nature of our planet.  Marine organisms produce much of the [[oxygen]] we breathe and probably help regulate the earth's [[climate]]. [[Shoreline]]s are in part shaped and protected by marine life, and some marine organisms even help create new land.
There are many practical reasons to study marine biology.  Marine life represents a vast resource, providing [[food]], [[medicine]], and raw materials, in addition to helping to support [[recreation]] and [[tourism]] all over the world. At a fundamental level, marine life helps determine the very nature of our planet.  Marine organisms produce much of the [[oxygen]] we breathe and probably help regulate the earth's [[climate]]. [[Shoreline]]s are in part shaped and protected by marine life, and some marine organisms even help create new land.


== Overview ==
== Overview ==
The species of organisms that are studied by marine biologists are extraordinarily diverse. In size, they range from the microscopic to the enormous. [[Genera]] that are too tiny to be seen without magnification include the  [[plankton]] and [[phytoplankton]], which can be as small as 0.02 [[micrometre | micrometers]], but are hugely important as the [[primary production | primary producers]] of the sea. The marine giants include  [[cetaceans]] (whales), which reach up to a reported 33 [[metre | meters]] (109 [[foot (unit of length) | feet]]) in length. The blue whale is the largest single living organism on earth in our time.


[[Image:Lionfish_in_coral_reef_2004-11-17.jpeg|thumb|right|Underwater life differs greatly from land life.]]
The salt-water habitats studied by marine biology are as diverse as the species, including everything from the tiny layers of surface water, in which organisms and abiotic items may be trapped in [[surface tension]] between the ocean and atmosphere, to the depths of the [[Ocean trench | abyssal trenches]], sometimes 10,000 meters or more beneath that surface, where darkness is constant and pressure so extreme that only creatures with special adaptations can survive. Marine habitats also include the [[coral reef]]s, [[kelp forest]]s, [[tidepools]], muddy, sandy, and rocky bottoms, and the open ocean (pelagic) zone, where solid objects are rare and the surface of the water is the only visible boundary.
 
The species of organisms that are studied by marine biologists are extraordinarily diverse. In size, they range from the microscopic to the enormous. [[Genera]] that are too tiny to be seen without magnification include the  [[plankton]] and [[phytoplankton]], which can be as small as 0.02 [[micrometre | micrometers]], but are hugely important as the [[primary production | primary producers]] of the sea. The marine giants include  [[cetaceans]] (whales), which reach up to a reported 33 [[metre | meters]] (109 [[foot (unit of length) | feet]]) in length. The blue whale is the largest single living organism on earth in our time.


The habitats studied by marine biology are as diverse as the species, including everything from the tiny layers of surface water in which organisms and abiotic items may be trapped in [[surface tension]] between the ocean and atmosphere, to the depths of the [[Ocean trench | abyssal trenches]], sometimes 10,000 meters or more beneath that surface, where darkness is constant and pressure so extreme that only creatures with special adaptations can survive even briefly. Marine habitats also include the [[coral reef]]s, [[kelp forest]]s, [[tidepools]], muddy, sandy, and rocky bottoms, and the open ocean (pelagic) zone, where solid objects are rare and the surface of the water is the only visible boundary.
Biologists agree that a large proportion of all life on [[Earth]] exists in the oceans, although exactly ''how'' large that proportion is remains unknown. While the oceans comprise about 71% of the Earth's surface, their great depths provide the marine habitat 300 times the volume of the terrestrial habitats on Earth.


A large proportion of all life on [[Earth]] exists in the oceansExactly how large the proportion is is still unknown.  While the oceans comprise about 71% of the Earth's surface, due to their depth they encompass about 300 times the habitable volume of the terrestrial habitats on Earth.
Many species are economically important to humans, including the [[fishery | food fishes]]. It is also becoming understood that marine ecology affects the land, and therefore influences the quality of human habitatsLife in the sea plays an important role in the maintenance of earth's mineral supplies (such as the [[carbon cycle]]), of air quality (such as [[General circulation model | Earth's respiration]], and in the movement of energy through the food chain in many ecosystems.


Many species are economically important to humans, including the [[fishery | food fishes]]. It is also becoming understood that the well-being of marine organisms and other organisms are linked in very fundamental ways.  Human understanding is growing of the relationship between life in the sea and important cycles such as that of matter (such as the [[carbon cycle]]), of air (such as [[General circulation model | Earth's respiration]], and movement of energy through ecosystems.  Large areas beneath the ocean surface still remain effectively unexplored.)
Large areas beneath the ocean surface still remain effectively unexplored.


== Subfields ==
== Subfields ==
The [[marine ecosystem]] is large, and thus there are many subfields of marine biology.  Most involve studying specializations of particular species (i.e., [[phycology]], [[invertebrate zoology]], [[ichthyology]]).
The [[marine ecosystem]] is large, and thus there are many subfields of marine biology.  Most involve studying specializations of particular species (i.e., [[phycology]], [[invertebrate zoology]], [[ichthyology]]).


Other subfields study the physical effects of continual immersion in [[sea water]] and the ocean in general, adaptation to a salty environment, and the affects of changing various oceanic properties on marine life.  A subfield of marine biology studies the relationships between oceans and ocean life, and global weather and environmental issues (such as [[carbon dioxide]] displacement).
Other subfields study the physical effects of continual immersion in [[sea water]] and the ocean in general, adaptation to a salty environment, and the affects of changing various oceanic properties on marine life.  A subfield of marine biology studies the relationships between oceans and ocean life, and global weather and [[Natural environment|environmental]] issues (such as [[carbon dioxide]] displacement).


Recent marine [[biotechnology]] has focused largely on [[marine biomolecule]]s, especially [[protein]]s, that may have uses in medicine or engineering. Marine environments are the home to many exotic biological materials that may inspire [[Bionics | biomimetic material]]s.
Recent marine [[biotechnology]] has focused largely on [[marine biomolecule]]s, especially [[protein]]s, that may have uses in medicine or engineering. Marine environments are the home to many exotic biological materials that may inspire [[Bionics | biomimetic material]]s.


An interesting branch of marine biology is [[aquaculture]]; which some countries do a lot of in the oceans, especially [[Japan]].
An interesting branch of marine biology is [[aquaculture]]; which some countries do a lot of in the oceans, especially Japan.


=== Related fields ===
=== Related fields ===
Marine biology is closely linked to both [[oceanography]] and [[biology]]. It also encompasses ideas from [[ecology]].  [[Fisheries science]] and [[marine conservation]] can be considered partial offshoots of marine biology.
Marine biology is closely linked to both [[oceanography]] and [[biology]]. It also encompasses ideas from [[ecology]].  [[Fisheries science]] and [[marine conservation]] can be considered partial offshoots of marine biology.


== Lifeforms ==
== Lifeforms ==
=== Microscopic life ===
=== Microscopic life ===
 
Microscopic life undersea is incredibly varied and still poorly understood.  For example, the role of [[virus]]es in marine ecosystems is barely being explored even in the beginning of the 21st century.
[[Image:Copepodkils.jpg|thumb|right|A [[copepod]].]]
 
Microscopic life undersea is incredibly varied and still poorly understood.  For example, the role of [[virus]]es in marine ecosystems is barely being explored even in the beginning of the [[21st century]].


The role of [[phytoplankton]] is better understood due to their critical position as the most numerous [[primary production | primary producers]] on Earth.  Phytoplankton are categorized into [[cyanobacteria]] (also called blue-green algae/bacteria), various types of [[algae]]: red, green, brown, and yellow-green, [[diatom]]s, [[dinoflagellate]]s, [[Euglenid | euglenoids]], [[coccolithophorid]]s, [[cryptomonad]]s, [[crysomonad]]s, [[chloromonad]]s, [[prasinomodad | prasinomonads]], and [[silicoflagellate]]s.
The role of [[phytoplankton]] is better understood due to their critical position as the most numerous [[primary production | primary producers]] on Earth.  Phytoplankton are categorized into [[cyanobacteria]] (also called blue-green algae/bacteria), various types of [[algae]]: red, green, brown, and yellow-green, [[diatom]]s, [[dinoflagellate]]s, [[Euglenid | euglenoids]], [[coccolithophorid]]s, [[cryptomonad]]s, [[crysomonad]]s, [[chloromonad]]s, [[prasinomodad | prasinomonads]], and [[silicoflagellate]]s.


[[Image:Giantkelp2_300.jpg|thumb|right|Giant [[kelp]].]]
[[Zooplankton]] tend to be somewhat larger, and not all are microscopic.  Many [[Protozoa]] are zooplankton, including dinoflagellates, [[Zooflagellate | zooflagellate]]s, [[Foraminifera | foraminiferans]], and [[radiolarian]]s.  Some of these (such as dinoflagellates) are also phytoplankton; the plant/animal distinction often breaks down in very small organisms.  Other zooplankton include [[cnidarian]]s, [[ctenophore]]s, [[chaetognatha | chaetognaths]], [[mollusc]]s, [[arthropod]]s, [[Tunicate | urochordates]], and [[annelid]]s such as [[polychaete]]s.  Many larger animals begin their life as zooplankton before they become large enough to take their familiar forms.  Two examples are fish [[larva]]e and sea stars (also called [[starfish]]).
 
[[Zooplankton]] tend to be somewhat larger, and not all are microscopic.  Many [[Protozoa]] are zooplankton, including dinoflagellates, [[Zooflagellate | zooflagellate]]s, [[Foraminifera | foraminiferans]], and [[radiolarian]]s.  Some of these (such as dinoflaggelates) are also phytoplankton; the plant/animal distinction often breaks down in very small organisms.  Other zooplankton include [[cnidarian]]s, [[ctenophore]]s, [[chaetognatha | chaetognaths]], [[mollusc]]s, [[arthropod]]s, [[Tunicate | urochordates]], and [[annelid]]s such as [[polychaete]]s.  Many larger animals begin their life as zooplankton before they become large enough to take their familiar forms.  Two examples are fish [[larva]]e and sea stars (also called [[starfish]]).


=== Plants and algae ===
=== Plants and algae ===
{{Image|Kelp.gif|right|200px|Kelp, ''Macrocystis pyrifera''<br><small>Photo credit: Michael Nishizaki</small>}}


Plant life is relatively rare undersea. Most of the [[niche]] occupied by plants on land is actually occupied by macroscopic [[algae]] in the ocean, such as ''[[Sargassum]]'' and [[kelp]] which are commonally known as [[seaweed]]s. The non algae plants that do survive in the sea are often found in shallow waters, such as the [[seagrass | seagrasses]] (examples of which are eelgrass, ''[[Zostera]]'', and turtlegrass, ''Thalassia''). These plants have adapted to the high salinity of the ocean environment. The [[foreshore | intertidal zone]] is also a good place to find plant life in the sea, where [[mangroves]] or [[cordgrass]] or [[beach grass]] might grow.
Most of the [[niche]]s occupied by plants on land are occupied by macroscopic [[algae]] in the ocean, which are commonly known as [[seaweed]]s. Most macroscopic  marine algae fall into one of three groups: 1). Rhodophyta ([[red algae]]); 2). Phaeophyta ([[brown algae]] including [[kelp]]) or; 3). Chlorophyta ([[green algae]]). Higher plants represent < 0.5% of the marine flora <ref name="Ackerman 1998">'''Ackerman, JD''' 1998. Is the limited diversity of higher plants in marine systems the result of biophysical limitations for reproduction or evolutionary and physiological constraint? ''Functional Ecology'' 12: 975–982.</ref> and are often found in shallow waters. All true marine plants are [[angiosperms]] and of [[polyphyletic]] origin (evolved separately in different places in the phylogenetic tree)<ref name="Les et al. 1997">'''Les, D.H., Cleland, M.A. & Waycott, M.A.''' 1997. Phylogenetic studies in Alismatidae. II. Evolution of marine angiosperms (seagrasses) and hydrophily? ''Systematic Botany'' 22: 443-463</ref>.


[[Image:Crown of Thorns-jonhanson.jpg|thumb|right|A [[crown-of-thorns starfish]].'']]
In the ocean, flowering plants are represented by the [[seagrass | seagrasses]] (examples of which are eelgrass, ''[[Zostera]]'', and turtlegrass, ''Thalassia''). These plants have adapted to the high salinity of the ocean environment. The [[foreshore | intertidal zone]] is also a good place to find plant life in the sea, where [[mangroves]] or [[cordgrass]] or [[beach grass]] might grow.


=== Marine invertebrates ===
=== Marine invertebrates ===
{{Image|Jellyfish.gif|right|200px|Jellyfish<br><small>Photo credit: Michael Nishizaki</small>}}


As on land, invertebrates make up a huge proportion of all life in the sea. Invertrebrate sea life includes Cnidaria such as [[Jellyfish]] and [[sea anemone]]; Ctenophora; sea [[worm]]s including [[phylum (biology) | phyla]]: Plathyhelminthes, Nemertea, Annelida, Sipuncula, Echiura, and the Phoronida;  Mollusca including [[shellfish]], [[squid]], [[octopus]];  [[Crustacean]]s;  Porifera including [[sponge]]s, [[Bryozoa]], [[Echinodermata]] including starfish; and [[Urochordata]] - [[sea squirt]]s or [[tunicate]]s.
As on land, invertebrates make up a huge proportion of all life in the sea. Invertebrate sea life includes Cnidaria such as [[Jellyfish]] and [[sea anemone]]; Ctenophora; sea [[worm]]s including [[phylum (biology) | phyla]]: Plathyhelminthes, Nemertea, Annelida, Sipuncula, Echiura, and the Phoronida;  Mollusca including [[shellfish]], [[squid]], [[octopus]];  [[Crustacean]]s including [[crab]]s, [[shrimp]] and [[barnacle]]s ;  Porifera including [[sponge]]s, [[Bryozoa]], [[Echinodermata]] including [[sea urchin]]s and [[sea star]]s; and [[Urochordata]] - [[sea squirt]]s or [[tunicate]]s.


=== Fish ===
=== Fish ===
{{Image|Fish-school.gif|right|200px|Shoaling fish<br><small>Photo credit: Michael Nishizaki</small>}}


[[Image:Carcharhinus longimanus 1.jpg |thumb|right|[[Oceanic whitetip shark]].]]
{{main | Fish}}
Fishes have [[evolution | evolved]] very different biological functions from other large organisms.  Fish anatomy includes a two-chambered heart, [[Operculum (Fish) | operculum]], secretory [[cell (biology) | cells]] that produce [[mucous]], [[swim bladder]], [[scale (zoology) | scales]], [[fin]]s, [[lip]]s and [[eye]]s.  Fish breathe by extracting [[oxygen]] from water through their gills.  Fins propel and stabilize the fish in the water.
Fishes have [[evolution | evolved]] very different biological functions from other large organisms.  Fish anatomy includes a two-chambered heart, [[Operculum (Fish) | operculum]], secretory [[cell (biology) | cells]] that produce [[mucous]], [[swim bladder]], [[scale (zoology) | scales]], [[fin]]s, [[lip]]s and [[eye]]s.  Fish breathe by extracting [[oxygen]] from water through their gills.  Fins propel and stabilize the fish in the water.


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=== Marine mammals ===
=== Marine mammals ===
[[Image:Sea_otter.jpg|thumb|right|A [[sea otter]].]]
{{main | Marine mammal}}
There are five main types of marine mammals.
There are five main types of marine mammals.


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== Reefs ==
== Reefs ==
[[Image:Assorted_living_corals.jpg|thumb|right|[[Coral]]s.]]
{{main | Coral reef}}
{{main | Coral reef}}


[[Reef]]s comprise some of the densest and most diverse habitats in the world.  The best-known types of reefs are [[tropical]] [[coral reef]]s which exist in most tropical waters; however, reefs can also exist in cold water.  Reefs are built up by [[coral]]s and other [[calcium]]-depositing animals, usually on top of a rocky outcrop on the ocean floor.  Reefs can also grow on other surfaces, which has made it possible to create [[artificial reef]]s. Coral reefs also support a huge community of life, including the corals themselves, their symbiotic [[zooxanthellae]], tropical fish and many other organisms.
[[Reef]]s comprise some of the densest and most diverse habitats in the world.  The best-known types of reefs are [[tropical]] [[coral reef]]s which exist in most tropical waters; however, reefs can also exist in cold water.  Reefs are built up by [[coral]]s and other [[calcium]]-depositing animals, usually on top of a rocky outcrop on the ocean floor.  Reefs can also grow on other surfaces, which has made it possible to create [[artificial reef]]s. Coral reefs also support a huge community of life, including the corals themselves, their symbiotic [[zooxanthellae]], tropical fish and many other organisms.


Much attention in marine biology is focused on coral reefs and the [[El Niño]] weather phenomenon. In [[1998]], coral reefs experienced a "once in a thousand years" bleaching event, in which vast expanses of reefs across the Earth died because sea surface temperatures rose well above normal.  Some reefs are recovering, but scientists say that 58[[%]] of the world's coral reefs are now endangered and predict that [[global warming]] could exacerbate this trend.
Much attention in marine biology is focused on coral reefs and the [[El Niño]] weather phenomenon. In 1998, coral reefs experienced a "once in a thousand years" bleaching event, in which vast expanses of reefs across the Earth died because sea surface temperatures rose well above normal.  Some reefs are recovering, but scientists say that 58[[%]] of the world's coral reefs are now endangered and predict that [[global warming]] could exacerbate this trend.


== Deep sea and trenches ==
== Deep sea and trenches ==


The ocean is deep, very deep in some places.  The deepest recorded measure to date is the [[Mariana Trench]], near the [[Philippines]], in the [[Pacific Ocean]] at 10924 m (35838 ft).  At such depths, [[water pressure]] is extreme and there is no sunlight, but some life still exists.  Small flounder (family [[Soleidae]]) fish and shrimp were seen by the American crew of the [[bathyscaphe]] ''[[Bathyscaphe Trieste | Trieste]]'' when it dove to the bottom in [[1960]].
The ocean is deep, very deep in some places.  The deepest recorded measure to date is the [[Mariana Trench]], near the [[Philippines]], in the [[Pacific Ocean]] at 10924 m (35838 ft).  At such depths, [[water pressure]] is extreme and there is no sunlight, but some life still exists.  Small flounder (family [[Soleidae]]) fish and shrimp were seen by the American crew of the [[bathyscaphe]] ''[[Bathyscaphe Trieste | Trieste]]'' when it dove to the bottom in 1960.


Other notable [[oceanic trench]]es include [[Monterey Canyon]], in the eastern Pacific, the [[Tonga Trench]] in the southwest at 10,882 m (35,702 feet), the [[Philippine Trench]], the [[Puerto Rico Trench]] at 8605 m (28232 ft), the [[Romanche Trench]] at 7760 m (24450 ft), [[Fram Basin]] in the [[Arctic Ocean]] at 4665 m (15305 ft), the [[Java Trench]] at 7450 m (24442 ft), and the [[South Sandwich Trench]] at 7235 m (23737 ft).
Other notable [[oceanic trench]]es include [[Monterey Canyon]], in the eastern Pacific, the [[Tonga Trench]] in the southwest at 10,882 m (35,702 feet), the [[Philippine Trench]], the [[Puerto Rico Trench]] at 8605 m (28232 ft), the [[Romanche Trench]] at 7760 m (24450 ft), [[Fram Basin]] in the [[Arctic Ocean]] at 4665 m (15305 ft), the [[Java Trench]] at 7450 m (24442 ft), and the [[South Sandwich Trench]] at 7235 m (23737 ft).
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In general, the deep sea is considered to start at the [[aphotic zone]], the point where sunlight loses its power of transference through the water.  Many life forms that live at these depths have the ability to create their own light.
In general, the deep sea is considered to start at the [[aphotic zone]], the point where sunlight loses its power of transference through the water.  Many life forms that live at these depths have the ability to create their own light.


Much life centers on [[seamount]]s that rise from the deeps, where fish and other sea life congregate to spawn and feed.  [[Hydrothermal vent]]s in the ocean floor act as [[oasis | oases]], as do their opposites, [[cold seeps]].  Such places support unique [[biome]]s and many new [[microbe]]s have been discovered at these sights.
Much life centers on [[seamount]]s that rise from the deeps, where fish and other sea life congregate to spawn and feed.  [[Hydrothermal vent]]s in the ocean floor act as [[oasis| oases]], as do their opposites, [[cold seeps]].  Such places support unique [[biome]]s and many new [[microbe]]s have been discovered at these sights.


Marine biologists are trying to amass a more complete mapping of underwater species with the help of modern techniques.  The deepest oceanic depressions are supposed to contain unknown species, possibly also of great potential interest for the theories on [[evolution]].
Marine biologists are trying to amass a more complete mapping of underwater species with the help of modern techniques.  The deepest oceanic depressions are supposed to contain unknown species, possibly also of great potential interest for the theories on [[evolution]].
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== How oceanic factors affect distribution of various organisms ==
== How oceanic factors affect distribution of various organisms ==


An active research topic in marine biology is to discover and map the [[biological life cycle | life cycle]]s of various species and where they spend their time. Marine  biologists study how the [[ocean current]]s, [[tide]]s and many other oceanic factors affect ocean lifeforms, including their growth, distribution and well-being.  This has only recently become technically feasible with advances in [[GPS]] and newer underwater visual devices.
An active research topic in marine biology is to describe the [[biological life cycle | life cycle]]s of various species and map where they spend their time. Marine  biologists study how [[ocean current]]s, [[tide]]s and many other oceanic factors affect ocean lifeforms, including their growth, distribution and well-being.  This has only recently become technically feasible with advances in [[GPS]] and newer underwater visual devices.
 
Most ocean life breeds in specific places, nests or not in others, spends time as juveniles in still others, and in maturity in yet others.  Scientists know little about where many species spent different parts of their life cycles.  For example, it is still largely unknown where [[sea turtle]]s travel.  Tracking devices do not work for some life forms, and the ocean is not friendly to [[technology]].  But these factors are being overcome in many instances.
 
== Famous marine biologists ==
 
[[Image:Jacques-Yves_Cousteau.jpg|right|thumb|Jacques-Yves Cousteau, co-inventer of the [[aqua-lung]], is well known for popularizing marine biology.]]
 
* Jakob Johan [[Adolf Appellöf]] ([[1857]]-[[1921]]), [[Sweden | Swedish]] marine zoologist.
* [[Joseph Ayers]], marine neurophysiologist and biomimetic researcher.
* [[Samuel Stillman Berry]] ([[1887]]-[[1984]]), U.S. marine zoologist.
* [[Henry Bryant Bigelow]] ([[1879]]–[[1967]]), U.S. marine biologist.
* [[Rachel Carson]] ([[1907]]-[[1964]]) American Marine Biologist and Author.
* [[Ilham Artüz]] ([[1924]]-[[1993]]) Turkish Marine Biologist and Oceanographer
* [[Carl Chun]] ([[1852]]-[[1914]]), [[Germany | German]] marine biologist
* [[Jacques-Yves Cousteau]] ([[1910]]-[[1997]]), [[France | French]] marine biologist and explorer.
* [[Anton Dohrn]] ([[1840]]-[[1909]]), German marine biologist.
* [[Sylvia Earle]] (born [[1935]]), American oceanographer.
* [[Johan Hjort]], Norwegian marine zoologist and one of the founders of [[ICES]].
* [[Bruno Hofer]] ([[1861]]-[[1916]]), German fisheries scientist.
* [[Hirohito]], Japanese emperor.
* [[August David Krohn]] ([[1803]]–[[1891]]), Russian/German zoologist.
* [[William Elford Leach]] ([[1790]]-[[1836]]), [[England | English]] zoologist and marine biologist.
* [[Philip S. Lobel]] Professor, [[Boston University]] Marine Program
* [[Nicholai Miklukho-Maklai]] ([[1846]]-[[1888]]), [[Russia | Russian]] marine biologist and anthropologist.
* [[John Murray (oceanographer) | Sir John Murray]] ([[1841]]-[[1914]]), [[Scotland | Scots]]-[[Canada | Canadian]] marine biologist.
* [[Harald Rosenthal]] (born [[1937]]), German hydrobiologist known for his work in fish farming and ecology.
* [[Michael Sars]] (1809–1869), [[Norway | Norwegian]] theologian and biologist.
* [[Georg Sars]] ([[1837]]-[[1927]]), Norwegian marine biologist.
* [[Ruth Turner]] ([[1915]]-[[2000]]), marine biologist.
* [[Robert Paine]] Professor, emeritus [[University of Washington]] intertidal Ecologist
* [[Charles Wyville Thompson]] ([[1832]]-[[1882]]), Scottish marine biologist
 
Source: [[List of biologists]].
 
== See also ==
 
* ''[[Aliens of the Deep]]''
* [[Animal]]
* [[Biology]]
* [[Fish]]
* [[The Life Aquatic with Steve Zissou]]
* [[List of publications in biology#Marine biology | Important publications in marine biology]]
* [[Mammals]]
* [[The Marine Mammal Center]]
* [[Ocean]]
* [[Plant]]
* [[Sea]]
 
== External links ==
 
* [http://www.whoi.edu/imageOfDay.do Oceanography Image of the Day], from the Woods Hole Oceanographic Institution
* [http://marinebio.org/ MarineBio: an evolving online tribute to Marine Biology and all Ocean life.]
* [http://www.mcsuk.org/ Marine Conservation Society: UK charity for seas and wildlife.]
* [http://www.seaturtle.org/tracking/?tag_id=49818 SeaTurtle.org: Tracking a sea turtle.]
* [http://www.marinebiology.co.uk/ Marinebiology.co.uk a marine biology resource.]
* [http://www.int-res.com/journals/meps/ Marine Ecology Progress Series (MEPS).]
* [http://www.mba.ac.uk/ Marine Biological Association of the United Kingdom.]
* [http://www.marlin.ac.uk/ Marine Life Information Network for Britain and Ireland - ''MarLIN''.]
* [http://content.lib.washington.edu/fishweb/index.html Freshwater and Marine Image Bank] The Freshwater and Marine Image Bank is an ongoing digital collection of images related to freshwater and marine topics. From the University of Washington Libraries Digital Images Collection
* [http://www.piscoweb.org/ PISCO - Partnership for Interdisciplinary Studies of Coastal Oceans]
* [http://www.meriresearch.org/ The Marine Environmental Research Institute]
* [http://www.oceanservice.noaa.gov/  The National Ocean Service]
* [[Wembury Marine Centre]]
 
{{Biology-footer}}


[[Category:Marine biology |Marine biology ]]
Many marine species occupy distinct habitats for breeding, spawning, nesting, juvenile development and adult dispersal.  Scientists know little about where many species spent different parts of their life cycles.  For example, it is still largely unknown where [[sea turtle]]s travel.  Tracking devices do not work for some life forms, and the ocean is not friendly to [[technology]].  But these factors are being overcome in many instances.


[[an:Biolochía marina]]
==References==
[[de:Meeresbiologie]]
<references/>[[Category:Suggestion Bot Tag]]
[[es:Biología marina]]
[[fr:Biologie marine]]
[[is:Sjávarlíffræði]]
[[lt:Jūrų biologija]]
[[nl:Mariene biologie]]
[[ja:海洋生物学]]
[[pt:Biologia marinha]]
[[simple:Marine biology]]
[[sv:Marinbiologi]]
[[tr:Deniz biyolojisi]]
[[zh:海洋生物学]]

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Marine biologists
Photo credit: Michael Nishizaki

Marine biology is the scientific study of the plants, animals and other organisms that live in the earth's ocean, seas, and related salt-water environments. Marine biologists are concerned not just with individual types of animals, but also with their inter-relationships with each other and their habitat.

There are many practical reasons to study marine biology. Marine life represents a vast resource, providing food, medicine, and raw materials, in addition to helping to support recreation and tourism all over the world. At a fundamental level, marine life helps determine the very nature of our planet. Marine organisms produce much of the oxygen we breathe and probably help regulate the earth's climate. Shorelines are in part shaped and protected by marine life, and some marine organisms even help create new land.

Overview

The species of organisms that are studied by marine biologists are extraordinarily diverse. In size, they range from the microscopic to the enormous. Genera that are too tiny to be seen without magnification include the plankton and phytoplankton, which can be as small as 0.02 micrometers, but are hugely important as the primary producers of the sea. The marine giants include cetaceans (whales), which reach up to a reported 33 meters (109 feet) in length. The blue whale is the largest single living organism on earth in our time.

The salt-water habitats studied by marine biology are as diverse as the species, including everything from the tiny layers of surface water, in which organisms and abiotic items may be trapped in surface tension between the ocean and atmosphere, to the depths of the abyssal trenches, sometimes 10,000 meters or more beneath that surface, where darkness is constant and pressure so extreme that only creatures with special adaptations can survive. Marine habitats also include the coral reefs, kelp forests, tidepools, muddy, sandy, and rocky bottoms, and the open ocean (pelagic) zone, where solid objects are rare and the surface of the water is the only visible boundary.

Biologists agree that a large proportion of all life on Earth exists in the oceans, although exactly how large that proportion is remains unknown. While the oceans comprise about 71% of the Earth's surface, their great depths provide the marine habitat 300 times the volume of the terrestrial habitats on Earth.

Many species are economically important to humans, including the food fishes. It is also becoming understood that marine ecology affects the land, and therefore influences the quality of human habitats. Life in the sea plays an important role in the maintenance of earth's mineral supplies (such as the carbon cycle), of air quality (such as Earth's respiration, and in the movement of energy through the food chain in many ecosystems.

Large areas beneath the ocean surface still remain effectively unexplored.

Subfields

The marine ecosystem is large, and thus there are many subfields of marine biology. Most involve studying specializations of particular species (i.e., phycology, invertebrate zoology, ichthyology).

Other subfields study the physical effects of continual immersion in sea water and the ocean in general, adaptation to a salty environment, and the affects of changing various oceanic properties on marine life. A subfield of marine biology studies the relationships between oceans and ocean life, and global weather and environmental issues (such as carbon dioxide displacement).

Recent marine biotechnology has focused largely on marine biomolecules, especially proteins, that may have uses in medicine or engineering. Marine environments are the home to many exotic biological materials that may inspire biomimetic materials.

An interesting branch of marine biology is aquaculture; which some countries do a lot of in the oceans, especially Japan.

Related fields

Marine biology is closely linked to both oceanography and biology. It also encompasses ideas from ecology. Fisheries science and marine conservation can be considered partial offshoots of marine biology.

Lifeforms

Microscopic life

Microscopic life undersea is incredibly varied and still poorly understood. For example, the role of viruses in marine ecosystems is barely being explored even in the beginning of the 21st century.

The role of phytoplankton is better understood due to their critical position as the most numerous primary producers on Earth. Phytoplankton are categorized into cyanobacteria (also called blue-green algae/bacteria), various types of algae: red, green, brown, and yellow-green, diatoms, dinoflagellates, euglenoids, coccolithophorids, cryptomonads, crysomonads, chloromonads, prasinomonads, and silicoflagellates.

Zooplankton tend to be somewhat larger, and not all are microscopic. Many Protozoa are zooplankton, including dinoflagellates, zooflagellates, foraminiferans, and radiolarians. Some of these (such as dinoflagellates) are also phytoplankton; the plant/animal distinction often breaks down in very small organisms. Other zooplankton include cnidarians, ctenophores, chaetognaths, molluscs, arthropods, urochordates, and annelids such as polychaetes. Many larger animals begin their life as zooplankton before they become large enough to take their familiar forms. Two examples are fish larvae and sea stars (also called starfish).

Plants and algae

Kelp, Macrocystis pyrifera
Photo credit: Michael Nishizaki

Most of the niches occupied by plants on land are occupied by macroscopic algae in the ocean, which are commonly known as seaweeds. Most macroscopic marine algae fall into one of three groups: 1). Rhodophyta (red algae); 2). Phaeophyta (brown algae including kelp) or; 3). Chlorophyta (green algae). Higher plants represent < 0.5% of the marine flora [1] and are often found in shallow waters. All true marine plants are angiosperms and of polyphyletic origin (evolved separately in different places in the phylogenetic tree)[2].

In the ocean, flowering plants are represented by the seagrasses (examples of which are eelgrass, Zostera, and turtlegrass, Thalassia). These plants have adapted to the high salinity of the ocean environment. The intertidal zone is also a good place to find plant life in the sea, where mangroves or cordgrass or beach grass might grow.

Marine invertebrates

Jellyfish
Photo credit: Michael Nishizaki

As on land, invertebrates make up a huge proportion of all life in the sea. Invertebrate sea life includes Cnidaria such as Jellyfish and sea anemone; Ctenophora; sea worms including phyla: Plathyhelminthes, Nemertea, Annelida, Sipuncula, Echiura, and the Phoronida; Mollusca including shellfish, squid, octopus; Crustaceans including crabs, shrimp and barnacles ; Porifera including sponges, Bryozoa, Echinodermata including sea urchins and sea stars; and Urochordata - sea squirts or tunicates.

Fish

Shoaling fish
Photo credit: Michael Nishizaki

Fishes have evolved very different biological functions from other large organisms. Fish anatomy includes a two-chambered heart, operculum, secretory cells that produce mucous, swim bladder, scales, fins, lips and eyes. Fish breathe by extracting oxygen from water through their gills. Fins propel and stabilize the fish in the water.

Well known fish include: sardines, anchovy, tuna, clownfish (also known as anemonefish), and bottom fish which include halibut and ling cod. Predators include sharks and barracuda.

Marine mammals

There are five main types of marine mammals.

Reefs

For more information, see: Coral reef.


Reefs comprise some of the densest and most diverse habitats in the world. The best-known types of reefs are tropical coral reefs which exist in most tropical waters; however, reefs can also exist in cold water. Reefs are built up by corals and other calcium-depositing animals, usually on top of a rocky outcrop on the ocean floor. Reefs can also grow on other surfaces, which has made it possible to create artificial reefs. Coral reefs also support a huge community of life, including the corals themselves, their symbiotic zooxanthellae, tropical fish and many other organisms.

Much attention in marine biology is focused on coral reefs and the El Niño weather phenomenon. In 1998, coral reefs experienced a "once in a thousand years" bleaching event, in which vast expanses of reefs across the Earth died because sea surface temperatures rose well above normal. Some reefs are recovering, but scientists say that 58% of the world's coral reefs are now endangered and predict that global warming could exacerbate this trend.

Deep sea and trenches

The ocean is deep, very deep in some places. The deepest recorded measure to date is the Mariana Trench, near the Philippines, in the Pacific Ocean at 10924 m (35838 ft). At such depths, water pressure is extreme and there is no sunlight, but some life still exists. Small flounder (family Soleidae) fish and shrimp were seen by the American crew of the bathyscaphe Trieste when it dove to the bottom in 1960.

Other notable oceanic trenches include Monterey Canyon, in the eastern Pacific, the Tonga Trench in the southwest at 10,882 m (35,702 feet), the Philippine Trench, the Puerto Rico Trench at 8605 m (28232 ft), the Romanche Trench at 7760 m (24450 ft), Fram Basin in the Arctic Ocean at 4665 m (15305 ft), the Java Trench at 7450 m (24442 ft), and the South Sandwich Trench at 7235 m (23737 ft).

In general, the deep sea is considered to start at the aphotic zone, the point where sunlight loses its power of transference through the water. Many life forms that live at these depths have the ability to create their own light.

Much life centers on seamounts that rise from the deeps, where fish and other sea life congregate to spawn and feed. Hydrothermal vents in the ocean floor act as oases, as do their opposites, cold seeps. Such places support unique biomes and many new microbes have been discovered at these sights.

Marine biologists are trying to amass a more complete mapping of underwater species with the help of modern techniques. The deepest oceanic depressions are supposed to contain unknown species, possibly also of great potential interest for the theories on evolution.

How oceanic factors affect distribution of various organisms

An active research topic in marine biology is to describe the life cycles of various species and map where they spend their time. Marine biologists study how ocean currents, tides and many other oceanic factors affect ocean lifeforms, including their growth, distribution and well-being. This has only recently become technically feasible with advances in GPS and newer underwater visual devices.

Many marine species occupy distinct habitats for breeding, spawning, nesting, juvenile development and adult dispersal. Scientists know little about where many species spent different parts of their life cycles. For example, it is still largely unknown where sea turtles travel. Tracking devices do not work for some life forms, and the ocean is not friendly to technology. But these factors are being overcome in many instances.

References

  1. Ackerman, JD 1998. Is the limited diversity of higher plants in marine systems the result of biophysical limitations for reproduction or evolutionary and physiological constraint? Functional Ecology 12: 975–982.
  2. Les, D.H., Cleland, M.A. & Waycott, M.A. 1997. Phylogenetic studies in Alismatidae. II. Evolution of marine angiosperms (seagrasses) and hydrophily? Systematic Botany 22: 443-463